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Are you using a CSS optimization plugin that may be incorrectly moving / modifying the inline CSS from WPSSO?
Perhaps you had a PHP issue when WPSSO created and compressed its CSS? If you haven’t already, you should define WP_DEBUG as true and make sure there are no PHP errors in the WordPress log. See here for more info: https://wpsso.com/docs/plugins/wpsso/installation/troubleshooting-guide/
To recreate the WPSSO compressed CSS, you can use the SSO > Clear Database Transients button.
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